Anoxygenic Phototrophs

The earliest photosynthesizing organisms were likely anoxy-genic phototrophs. These use hydrogen sulfide or organic compounds rather than water as a source of electrons when making reducing power in the form of NADPH, and therefore they do not generate O2. Modern anoxygenic phototrophs are a phylogenetically diverse group of bacteria that inhabit a restricted ecological niche that provides adequate light penetration yet little or no O2; most often, they are found in aquatic habitats such as bogs, lakes, and the upper layer of muds. ■ reducing power, p. 134

The photosynthetic systems of the anoxygenic phototrophs are fundamentally different from those of plants, algae, and cyanobacteria. They have a unique type of chlorophyll called bacteriochlorophyll. This pigment absorbs wavelengths of light that penetrate to greater depths and are not used by other photosynthetic organisms. ■ photosynthetic pigments, p. 156

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